This signal box was west of Greenhill Lower Junction and Bonnybridge Junction on the Caledonian Railway.
This timber box opened in 1886 to replace Dalnair box, further west at the start of the Greenhill loops, and the previous Greenhill South box, a little further east by the west end of Bonnybridge Junction Yard.
The 1886 box was located on the north side of the line. It remained in use until 1975 when it was taken over by Greenhill Upper Junction box during a resignalling and the layout was simplified, having been rationalised previously.